I think, but double check it, the 4 stud wheels were the same pattern as VW Beetle (the original, not this "modern" Golf based thingy...).
I'll guarantee it! Among the many wheels I sold Woodstock (a 72 4/4 with bolt on wheels) with were a set of wheels with VW beetle centres that I had Motor Wheel Services put new rims on.
The wheelnuts for the rostyle ones would have fitted, but so did the wheelnuts for an LT35 van. And they were smaller so less weight.