It is often said they self tighten. But given the wheels are on splines I cant see how. There is a bit of circular momentum from the ears... but that acts in opposite direction when accelerating and braking... I suppose the braking force is higher than the accelerating force which would net out as a tightening force. But the mass of the ears is pretty small I would have thought it had minimal effect?
By the way, this is an innocent question.... not got an axe to grind