All cars have ‘souls’ mainly because of the necessity to have two headlights which infer eyes
The radiator as a mostly angry or aggressive mouth is quite a recent design meme that could only really developed after the flat radiator and first appeared in the very late 40’s but boomed in 50’s sports cars
The earliest ‘mouth’ grills I can bring readily to mind of are from about the late 40’s - Aston Martin DB1 and a Ferrari or two (I am sure there are more)
Before then the headlights were still eyes but the grills were more noses as they were higher and more squashed
In America the early 50’s, the Corvette comes to mind as a good example of an aggressive mouth (it even has teeth) so it probably moved in tandem with a lower bonnet and lower wider radiator in sports cars
Now with SUV’s, designers have raised the front end but tried to keep the agressive mouth of a sport car and made a super angry screaming mouth of a grill
