They changed the louvres with the launch of the first Roadster series1. Only that car had an intake directly at the inside bonnet of the louvres. My unimportant guess: MMC did some survey at that time, perhaps the airflow was better by changing the direction of the louvres. Then the following series had a different intake but now the tools for the bonnet had been altered and will still be used until today.
Thanks Heinz, you've reminded me that the first time I saw inverted louvres was on my 2005 roadster mk1. Mine had no inner fitting to the louvres as some others had. The thing is though, have any tests of airflow under there been made since the change? Nick