They changed the louvres with the launch of the first Roadster series1. Only that car had an intake directly at the inside bonnet of the louvres. My unimportant guess: MMC did some survey at that time, perhaps the airflow was better by changing the direction of the louvres. Then the following series had a different intake but now the tools for the bonnet had been altered and will still be used until today.
Funny thing. We had an ex bonnet maker here as a guest between Christmas and new year. I happened to mention the direction of the bonnet louvres in conversation about my bonnets and I had thought that it was to do with safety but he told me that it was changed specifically to accommodate the air intake on the first Roadsters. So there is your answer.